Contact

Robert K. Colwell

Museum Curator Adjoint in Entomology


robertkcolwell [at] gmail.com


Museum of Natural History

University of Colorado

Boulder, CO 80309, USA




robertkcolwell [at] gmail.com


Museum of Natural History

University of Colorado

Boulder, CO 80309, USA



Publications


Repeated evolution of unorthodox feeding styles drives a negative correlation between foot size and bill length in hummingbirds


Robert K. Colwell, Thiago F. Rangel, Karolina Fučiková, Diego Sustaita, Gregor M. Yanega, Alejandro Rico-Guevara

The American Naturalist, [Just accepted preprint]



Monitoring recovery of tree diversity during tropical forest restoration: lessons from long-term trajectories of natural regeneration


Robin Chazdon, Natalia Norden, Robert K Colwell, Anne Chao

Royal Society of London. Philosophical Transactions B. Biological Sciences, The Royal Society Publishing, 2022



Language and ethnobiological skills decline precipitously in Papua New Guinea, the world’s most linguistically diverse nation


A. Kik, M. Adamec, A. Aikhenvald, Jarmila Bajzekova, Nigel Baro, Claire Bowern, R. K. Colwell, P. Drozd, P. Duda, Sentiko Ibalim, L. Jorge, J. Mogina, Ben Ruli, K. Sam, Hannah S. Sarvasy, S. Saulei, G. Weiblen, J. Zrzavý, V. Novotný

Proceedings of the National Academy of Sciences, 2021



Landscape dynamics promoted the evolution of mega-diversity in South American freshwater fishes


F. Cassemiro, J. Albert, A. Antonelli, A. Menegotto, R. Wüest, M. T. Coelho, D. Bailly, Valéria F. B. da Silva, A. Frota, W. J. da Graça, Reginaldo Ré, T. Ramos, Anielly Galego de Oliveira, M. S. Dias, R. K. Colwell, T. Rangel, C. Graham

2021



Demographic shifts, inter-group contact, and environmental conditions drive language extinction and diversification


Marco Tulio Pacheco Coelho1, H. Haynie, Claire Bowern, R. K. Colwell, Simon J. Greenhill, K. Kirby, T. Rangel, M. Gavin

2021



Quantifying sample completeness and comparing diversities among assemblages


A. Chao, Y. Kubota, D. Zelený, Chun‐Huo Chiu, Ching‐Feng Li, B. Kusumoto, M. Yasuhara, S. Thorn, Chih‐Lin Wei, Mark John Costello, R. K. Colwell

2020



Alpine archaeology of Alta Toquima and the Mt. Jefferson Tablelands (Nevada) : the archaeology of Monitor Valley, contribution 4.


D. Thomas, J. Bean, Gregory R. Burns, T. Canaday, D. Charlet, R. K. Colwell, B. Culleton, J. Eerkens, Nicholas P. Freeland, D. Graybill, D. Grayson, T. Harper, R. Hughes, J. Jiménez, D. Kennett, C. Millar, A. Novick, L. Pendleton, A. Rankin, David Rhode, J. Rosenthal, I. Rovner, H. Spero, Nathan E. Stevens

2020



Building mountain biodiversity: Geological and evolutionary processes


C. Rahbek, M. K. Borregaard, A. Antonelli, R. K. Colwell, Ben G Holt, D. Nogues‐Bravo, C. Rasmussen, K. Richardson, M. Rosing, R. Whittaker, J. Fjeldså

Science, 2019



Humboldt’s enigma: What causes global patterns of mountain biodiversity?


C. Rahbek, M. K. Borregaard, R. K. Colwell, B. Dalsgaard, Ben G Holt, N. Morueta‐Holme, D. Nogues‐Bravo, R. Whittaker, J. Fjeldså

Science, 2019



The distributions of morphologically specialized hummingbirds coincide with floral trait matching across an Andean elevational gradient


Jesper Sonne, Thais B. Zanata, Ana M. Martín González, Nixon L. Cumbicus Torres, J. Fjeldså, R. K. Colwell, Boris A. Tinoco, C. Rahbek, B. Dalsgaard

Biotropica, 2019



Drivers of geographical patterns of North American language diversity


Marco Túlio Pacheco Coelho, E. Pereira, H. Haynie, T. Rangel, P. Kavanagh, K. Kirby, S. Greenhill, Claire Bowern, R. Gray, R. K. Colwell, N. Evans, M. Gavin

Proceedings of the Royal Society B, 2019



The distributions of morphologically specialized hummingbirds coincidewith floral trait matching across an Andean elevational gradient


Jesper Sonne, Thais B Zanata, Ana M Mart{\'\i}n Gonz{\'a}lez, Nixon L Cumbicus Torres, Jon Fjelds{\aa}, Robert K Colwell, Boris A Tinoco, Carsten Rahbek, Bo Dalsgaard

2019



Managing consequences of climate‐driven species redistribution requires integration of ecology, conservation and social science


T. Bonebrake, Christopher J. Brown, J. Bell, J. Blanchard, A. Chauvenet, C. Champion, I. Chen, T. Clark, R. K. Colwell, F. Danielsen, A. Dell, J. Donelson, B. Evengård, S. Ferrier, S. Frusher, R. A. Garcia, R. Griffis, A. Hobday, Marta A. Jarzyna, Emma Lee, J. Lenoir, Hlif I. Linnetved, Victoria Y. Martin, Phillipa C. McCormack, J. McDonald, E. McDonald‐Madden, N. Mitchell, T. Mustonen, J. Pandolfi, N. Pettorelli, H. Possingham, P. Pulsifer, M. Reynolds, Brett R. Scheffers, Cascade J. B. Sorte, J. Strugnell, M. Tuanmu, S. Twiname, A. Vergés, Cecilia Villanueva, E. Wapstra, T. Wernberg, G. Pecl

Biological reviews of the Cambridge Philosophical Society, 2018



Spatially explicit paleoclimate reconstruction of South America for the last 800 thousand years


T. Rangel, N. Edwards, P. Holden, J. Diniz-Filho, W. Gosling, M. T. Coelho, F. Cassemiro, C. Rahbek, R. K. Colwell

2018



Modeling the ecology and evolution of biodiversity: Biogeographical cradles, museums, and graves


T. Rangel, N. Edwards, P. Holden, J. Diniz-Filho, W. Gosling, M. T. Coelho, F. Cassemiro, C. Rahbek, R. K. Colwell

Science, 2018



Response to Qian et al. (2017): Daily and seasonal climate variations are both critical in the evolution of species’ elevational range size


Wei-ping Chan, I. Chen, R. K. Colwell, Wei-Chung Liu, Cho-ying Huang, S. Shen

Journal of Biogeography, 2018



Deciphering the enigma of undetected species, phylogenetic, and functional diversity based on Good-Turing theory.


A. Chao, Chun‐Huo Chiu, R. K. Colwell, Luiz F. S. Magnago, R. Chazdon, N. Gotelli

Ecology, 2017



Environment-induced changes in selective constraints on social learning during the peopling of the Americas


Briggs Buchanan, A. Chao, Chun‐Huo Chiu, R. K. Colwell, M. J. O’Brien, Angelia Werner, M. Eren

Scientific reports, 2017



Biodiversity redistribution under climate change: Impacts on ecosystems and human well-being


G. Pecl, M. Araújo, J. Bell, J. Blanchard, T. Bonebrake, I. Chen, T. Clark, R. K. Colwell, F. Danielsen, B. Evengård, L. Falconi, S. Ferrier, S. Frusher, R. A. Garcia, R. Griffis, A. Hobday, C. Janion‐Scheepers, Marta A. Jarzyna, S. Jennings, J. Lenoir, Hlif I. Linnetved, Victoria Y. Martin, Phillipa C. McCormack, J. McDonald, N. Mitchell, T. Mustonen, J. Pandolfi, N. Pettorelli, E. Popova, S. Robinson, Brett R. Scheffers, J. Shaw, Cascade J. B. Sorte, J. Strugnell, J. Sunday, M. Tuanmu, A. Vergés, Cecilia Villanueva, T. Wernberg, E. Wapstra, Stephen E. Williams

Science, 2017



Elevational species richness gradients in a hyperdiverse insect taxon: a global meta-study on geometrid moths


Jan Beck, C. McCain, J. Axmacher, L. Ashton, F. Bärtschi, G. Brehm, Sei‐Woong Choi, O. Čížek, R. K. Colwell, K. Fiedler, C. Francois, Steven A. Highland, J. Holloway, J. Intachat, T. Kadlec, R. Kitching, S. Maunsell, T. Merckx, A. Nakamura, E. Odell, W. Sang, P. Toko, J. Zámečník, Y. Zou, V. Novotný

2017



Process-based modelling shows how climate and demography shape language diversity


M. Gavin, T. Rangel, Claire Bowern, R. K. Colwell, K. Kirby, C. Botero, M. Dunn, R. Dunn, J. McCarter, M. T. Coelho, R. Gray

2017



Mechanism, Process, and Causation in Ecological Models: A Reply to McGill and Potochnik.


S. Connolly, S. Keith, R. K. Colwell, C. Rahbek

Trends in ecology & evolution, 2017



Opinion Process , Mechanism , and Modeling in Macroecology


S. Connolly, S. Keith, R. K. Colwell, C. Rahbek

2017



Process, Mechanism, and Modeling in Macroecology.


S. Connolly, S. Keith, R. K. Colwell, C. Rahbek

Trends in ecology & evolution, 2017



What Makes Models Mechanistic or Process-Based ?


S. Connolly, S. Keith, R. K. Colwell, C. Rahbek

2017



Turning Up the Heat on a Hotspot: DNA Barcodes Reveal 80% More Species of Geometrid Moths along an Andean Elevational Gradient


G. Brehm, P. Hebert, R. K. Colwell, Marc-Oliver Adams, Florian Bodner, Katrin Friedemann, Lars Möckel, K. Fiedler

PloS one, 2016



Statistical Analysis of Paradigmatic Class Richness Supports Greater Paleoindian Projectile-Point Diversity in the Southeast


M. Eren, A. Chao, Chun‐Huo Chiu, R. K. Colwell, Briggs Buchanan, Matthew Boulanger, John Darwent, M. J. O’Brien

American Antiquity, 2016



Appendix G. A user's guide for program CLAM, with instructions and a link for downloading the program.


R. Chazdon, A. Chao, R. K. Colwell, Shang-Yi Lin, Natalia Norden, Susan G. Letcher, David B. Clark, B. Finegan, J. P. Arroyo

2016



Appendix A. Description of vegetation inventory data sets used in this study.


R. Chazdon, A. Chao, R. K. Colwell, Shang-Yi Lin, Natalia Norden, Susan G. Letcher, David B. Clark, B. Finegan, J. P. Arroyo

2016



Midpoint attractors and species richness: Modelling the interaction between environmental drivers and geometric constraints.


R. K. Colwell, N. Gotelli, L. Ashton, Jan Beck, G. Brehm, T. Fayle, K. Fiedler, M. Forister, M. Kessler, R. Kitching, P. Klimes, Jürgen Kluge, J. Longino, S. Maunsell, C. McCain, Jimmy Moses, Sarah Noben, K. Sam, L. Sam, A. Shapiro, Xiangping Wang, V. Novotný

Ecology letters, 2016



Seasonal and daily climate variation have opposite effects on species elevational range size


Wei-ping Chan, I. Chen, R. K. Colwell, Wei-Chung Liu, Cho-ying Huang, S. Shen

Science, 2016



An estimate of the number of tropical tree species


J. Slik, V. Arroyo‐Rodríguez, S. Aiba, P. Álvarez-Loayza, L. Alves, P. Ashton, P. Balvanera, M. Bastian, P. Bellingham, E. van den Berg, L. Bernacci, Polyanna da Conceição Bispo, L. Blanc, K. Böhning‐Gaese, P. Boeckx, F. Bongers, B. Boyle, M. Bradford, F. Brearley, Mireille Breuer-Ndoundou Hockemba, S. Bunyavejchewin, Darley Calderado Leal Matos, M. Castillo-Santiago, E. Catharino, Shauna-Lee Chai, Yukai Chen, R. K. Colwell, R. Chazdon, C. Clark, D. Clark, D. Clark, H. Culmsee, K. Damas, H. Dattaraja, G. Dauby, P. Davidar, S. DeWalt, J. Doucet, Á. Duque, G. Durigan, K. Eichhorn, P. V. Eisenlohr, E. Eler, C. Ewango, Nina Farwig, K. Feeley, L. Ferreira, Richard Field, Ary T. de Oliveira Filho, C. Fletcher, O. Forshed, G. Franco, G. Fredriksson, T. Gillespie, J. Gillet, G. Amarnath, Daniel M. Griffith, J. Grogan, N. Gunatilleke, David Harris, R. Harrison, A. Hector, J. Homeier, N. Imai, A. Itoh, P. Jansen, C. Joly, B. D. De Jong, K. Kartawinata, E. Kearsley, D. Kelly, D. Kenfack, M. Kessler, K. Kitayama, R. Kooyman, E. Larney, Y. Laumonier, S. Laurance, W. Laurance, M. Lawes, I. Amaral, Susan G. Letcher, J. Lindsell, Xinghui Lu, A. Mansor, A. Marjokorpi, Emanuel H. Martin, H. Meilby, F. Melo, D. Metcalfe, V. Medjibe, J. Metzger, J. Millet, D. Mohandass, J. Montero, M. de Morisson Valeriano, B. Mugerwa, H. Nagamasu, R. Nilus, S. Ochoa-Gaona, Onrizal, N. Page, P. Parolin, M. Parren, N. Parthasarathy, E. Paudel, Andrea Permana, M. Piedade, N. Pitman, L. Poorter, A. Poulsen, J. Poulsen, J. Powers, R. Prasad, J. Puyravaud, J. Razafimahaimodison, J. Reitsma, J. dos Santos, Wilson Roberto Spironello, Hugo Romero-Saltos, F. Rovero, A. Rozak, K. Ruokolainen, E. Rutishauser, F. Saiter, P. Saner, Bráulio A. Santos, Fernanda Santos, S. K. Sarker, Manichanh Satdichanh, C. Schmitt, J. Schöngart, M. Schulze, M. Suganuma, D. Sheil, Eduardo da Silva Pinheiro, P. Sist, T. Stévart, R. Sukumar, I. Sun, T. Sunderland, H. Suresh, E. Suzuki, M. Tabarelli, Jangwei Tang, N. Targhetta, I. Theilade, Duncan W. Thomas, P. Tchouto, Johanna Hurtado, R. Valencia, J. V. van Valkenburg, Tran Van Do, R. Vásquez, H. Verbeeck, V. Adekunle, S. Vieira, C. Webb, Timothy J. S. Whitfeld, S. Wich, John N. Williams, F. Wittmann, Hannsjoerg Wöll, Xiaobo Yang, C. Y. Adou Yao, S. Yap, T. Yoneda, R. Zahawi, R. Zakaria, R. Zang, R. L. de Assis, Bruno Garcia Luize, Eduardo M. Venticinque

Proceedings of the National Academy of Sciences, 2015



Phylogenetic uncertainty revisited: Implications for ecological analyses


T. Rangel, R. K. Colwell, G. Graves, K. Fučíková, C. Rahbek, J. Diniz-Filho

Evolution; international journal of organic evolution, 2015



Defining and observing stages of climate-mediated range shifts in marine systems


A. Bates, G. Pecl, S. Frusher, A. Hobday, T. Wernberg, D. Smale, J. Sunday, N. Hill, N. Dulvy, R. K. Colwell, N. Holbrook, E. Fulton, D. Slawinski, M. Feng, G. Edgar, B. Radford, P. Thompson, R. Watson

2014



Rarefaction and extrapolation with Hill numbers: a framework for sampling and estimation in species diversity studies


A. Chao, N. Gotelli, T. Hsieh, Elizabeth L. Sander, K. Ma, R. K. Colwell, A. Ellison

2014



How Ants Drop Out: Ant Abundance on Tropical Mountains


J. Longino, M. Branstetter, R. K. Colwell

PloS one, 2014



Thermal-safety margins and the necessity of thermoregulatory behavior across latitude and elevation


J. Sunday, A. Bates, M. Kearney, R. K. Colwell, N. Dulvy, J. Longino, R. Huey

Proceedings of the National Academy of Sciences, 2014



Understanding historical and current patterns of species richness of babblers along a 5000‐m subtropical elevational gradient


Yongjie Wu, R. K. Colwell, Naijian Han, Ruiying Zhang, Wenjuan Wang, Q. Quan, Chunlan Zhang, G. Song, Y. Qu, F. Lei

2014



Quantifying temporal change in biodiversity: challenges and opportunities


M. Dornelas, A. Magurran, S. T. Buckland, A. Chao, R. Chazdon, R. K. Colwell, T. Curtis, K. Gaston, N. Gotelli, M. Kosnik, B. McGill, J. L. McCune, H. Morlon, P. Mumby, L. Øvreås, A. Studeny, M. Vellend

Proceedings of the Royal Society B: Biological Sciences, 2013



Toward a Mechanistic Understanding of Linguistic Diversity


M. Gavin, C. Botero, Claire Bowern, R. K. Colwell, M. Dunn, R. Dunn, R. Gray, K. Kirby, J. McCarter, Adam Powell, T. Rangel, J. Stepp, Michelle Trautwein, J. Verdolin, G. Yanega

2013



Explaining the species richness of birds along a subtropical elevational gradient in the Hengduan Mountains


Yongjie Wu, R. K. Colwell, C. Rahbek, Chunlan Zhang, Q. Quan, Changke Wang, F. Lei

2013



Report on Workshop on Possible Ecological and Evolutionary Impacts of Bioengineered Organisms Released into the Environment


James H. Brown, R. K. Colwell, R. Lenski, B. Levin, MonteLloyd, P. Regal, DanielSimberloff

2012



Specimen-based modeling, stopping rules, and the extinction of the Ivory-billed Woodpecker.


N. Gotelli, A. Chao, R. K. Colwell, W. Hwang, G. Graves

Conservation biology : the journal of the Society for Conservation Biology, 2012



Biodiversity: Species loss revisited


C. Rahbek, R. K. Colwell

Nature, 2011



A novel statistical method for classifying habitat generalists and specialists.


R. Chazdon, A. Chao, R. K. Colwell, Shang-Yi Lin, Natalia Norden, Susan G. Letcher, D. Clark, B. Finegan, J. P. Arroyo

Ecology, 2011



Estimating species richness


Nicholas J Gotelli, Robert K Colwell

Biological diversity: frontiers in measurement and assessment. A. E. Magurran and B. J. McGill, eds., Oxford University Press, Oxford, UK, 2010, pp. 39--54



A stochastic, evolutionary model for range shifts and richness on tropical elevational gradients under Quaternary glacial cycles


R. K. Colwell, T. Rangel

Philosophical Transactions of the Royal Society B: Biological Sciences, 2010



Sufficient sampling for asymptotic minimum species richness estimators


Anne Chao, Robert K Colwell, Chih-Wei Lin, Nicholas J Gotelli

Ecology, vol. 90, Ecological Society of America, 2009, pp. 1125--1133



EstimateS 8.2. 0.--Statistical estimation of species richness and shared species from samples


RK Colwell

User’s Guide and Application. Department of Ecology and Evolutionary Biology, University of Connecticut, Storrs, 2009



Lawrence B. Slobodkin


Robert K Colwell, Douglas J Futuyma

EVOLUTIONARY ECOLOGY LTD UNIV ARIZONA, 321 BIOSCIENCES WEST, TUCSON, AZ~…, 2009



Peaks, plateaus, canyons, and craters: the complex geometry of simple mid-domain effect models


Robert K Colwell, Nicholas J Gotelli, Carsten Rahbek, Gary L Entsminger, Catherine Farrell, Gary R Graves

Evolutionary Ecology Research, 2009



Peaks, plateaus, canyons, and craters: the complex geometry of simple mid-domain effect models


Robert K Colwell, Nicholas J Gotelli, Carsten Rahbek, Gary L Entsminger, Catherine Farrell, Gary R Graves

Evolutionary Ecology Research, 2009



Biodiversity: concepts, patterns, and measurement


Robert K Colwell, others

The Princeton guide to ecology, vol. 663, Princeton University Press Princeton, 2009, pp. 257--263



Hutchinson's duality: the once and future niche


Robert K Colwell, Thiago F Rangel

Proceedings of the National Academy of Sciences, vol. 106, National Academy of Sciences, 2009, pp. 19651--19658



The sixth mass coextinction: are most endangered species parasites and mutualists?


Robert R Dunn, Nyeema C Harris, Robert K Colwell, Lian Pin Koh, Navjot S Sodhi

Proceedings of the Royal Society B: Biological Sciences, vol. 276, The Royal Society, 2009, pp. 3037--3045



Elevational patterns of diversity and abundance of eusocial paper wasps (Vespidae) in Costa Rica


Anjali Kumar, John T Longino, Robert K Colwell, Sean O'Donnell

Biotropica, vol. 41, Blackwell Publishing Inc Malden, USA, 2009, pp. 338--346



Vulnerability and resilience of tropical forest species to land-use change


Nigel E Stork, Jonathan A Coddington, Robert K Colwell, Robin L Chazdon, Christopher W Dick, Carlos A Peres, Sean Sloan, Kathy Willis

Conservation Biology, vol. 23, Blackwell Publishing Inc, 2009, pp. 1438--1447



Vulnerability and resilience of tropical forest species to land-use change


Nigel E Stork, Jonathan A Coddington, Robert K Colwell, Robin L Chazdon, Christopher W Dick, Carlos A Peres, Sean Sloan, Kathy Willis

Conservation Biology, vol. 23, Blackwell Publishing Inc, 2009, pp. 1438--1447



Distribution of megabenthic gastropods along environmental gradients: the mid-domain effect and beyond


Alvar Carranza, Robert K Colwell, Thiago Fernando LVB Rangel

Marine Ecology Progress Series, vol. 367, 2008, pp. 193--202



Global warming, elevational range shifts, and lowland biotic attrition in the wet tropics


Robert K Colwell, Gunnar Brehm, Catherine L Cardel{\'u}s, Alex C Gilman, John T Longino

science, vol. 322, American Association for the Advancement of Science, 2008, pp. 258--261



Correlates of extinction proneness in tropical angiosperms


Navjot S Sodhi, Lian Pin Koh, Kelvin S-H Peh, Hugh TW Tan, Robin L Chazdon, Richard T Corlett, Tien Ming Lee, Robert K Colwell, Barry W Brook, Cagan H Sekercioglu, others

Diversity and Distributions, vol. 14, Blackwell Publishing Ltd Oxford, UK, 2008, pp. 1--10



The role of environment and mid-domain effect on moth species richness along a tropical elevational gradient


Gunnar Brehm, Robert K Colwell, J{\"u}rgen Kluge

Global Ecology and Biogeography, vol. 16, Blackwell Publishing Ltd Oxford, UK, 2007, pp. 205--219



A comparison of taxon co-occurrence patterns for macro- and microorganisms


M Claire Horner-Devine, Jessica M Silver, Mathew A Leibold, Brendan JM Bohannan, Robert K Colwell, Jed A Fuhrman, Jessica L Green, Cheryl R Kuske, Jennifer BH Martiny, Gerard Muyzer, others

Ecology, vol. 88, 2007, pp. 1345--1353



Predicting the ecosystem consequences of biodiversity loss: the biomerge framework


Shahid Naeem, Robert Colwell, Sandra D{\'\i}az, Jennifer Hughes, Claire Jouseau, Sandra Lavorel, Peter Morin, Owen Petchey, Justin Wright, others

Terrestrial ecosystems in a changing world, Springer, Berlin, Heidelberg, 2007, pp. 113--126



Predicting continental-scale patterns of bird species richness with spatially explicit models


Carsten Rahbek, Nicholas J Gotelli, Robert K Colwell, Gary L Entsminger, Thiago Fernando LVB Rangel, Gary R Graves

Proceedings of the Royal Society B: Biological Sciences, vol. 274, The Royal Society London, 2007, pp. 165--174



Species richness and evolutionary niche dynamics: a spatial pattern--oriented simulation experiment


Thiago Fernando LVB Rangel, Jos{\'e} Alexandre F Diniz-Filho, Robert K Colwell

The American Naturalist, vol. 170, The University of Chicago Press, 2007, pp. 602--616



Vascular epiphyte distribution patterns: explaining the mid-elevation richness peak


Catherine L Cardel{\'u}s, Robert K Colwell, James E Watkins Jr

Journal of Ecology, British Ecological Society, 2006, pp. 144--156



Abundance-based similarity indices and their estimation when there are unseen species in samples


Anne Chao, Robin L Chazdon, Robert K Colwell, Tsung-Jen Shen

Biometrics, vol. 62, Blackwell Publishing Inc, 2006, pp. 361--371



The river domain: why are there more species halfway up the river?


Robert R Dunn, Robert K Colwell, Christer Nilsson

Ecography, vol. 29, Blackwell Publishing Ltd Copenhagen, 2006, pp. 251--259



Microbial biogeography: putting microorganisms on the map


Jennifer B Hughes Martiny, Brendan JM Bohannan, James H Brown, Robert K Colwell, Jed A Fuhrman, Jessica L Green, M Claire Horner-Devine, Matthew Kane, Jennifer Adams Krumins, Cheryl R Kuske, others

Nature Reviews Microbiology, vol. 4, Nature Publishing Group, 2006, pp. 102--112



Species richness and distribution of ferns along an elevational gradient in Costa Rica


James E Watkins Jr, Catherine Cardel{\'u}s, Robert K Colwell, Robbin C Moran

American Journal of Botany, vol. 93, Botanical Society of America, 2006, pp. 73--83



Species loss and aboveground carbon storage in a tropical forest


Daniel E Bunker, Fabrice DeClerck, Jason C Bradford, Robert K Colwell, Ivette Perfecto, Oliver L Phillips, Mahesh Sankaran, Shahid Naeem

Science, vol. 310, American Association for the Advancement of Science, 2005, pp. 1029--1031



A new statistical approach for assessing similarity of species composition with incidence and abundance data


Anne Chao, Robin L Chazdon, Robert K Colwell, Tsung-Jen Shen

Ecology letters, vol. 8, Blackwell Science Ltd Oxford, UK, 2005, pp. 148--159



A new statistical approach for assessing similarity of species composition with incidence and abundance data


Anne Chao, Robin L Chazdon, Robert K Colwell, Tsung-Jen Shen

Ecology letters, vol. 8, Blackwell Science Ltd Oxford, UK, 2005, pp. 148--159



Estimation of species richness at large spatial scales: Potential, procedures, pitfalls.


Robert K Colwell

A. Bergamini, Ed. Landscape scale biodiversity assessment, the problem of scaling, European Platform for Biodiversity Research Strategy (EPBRS), Budapest, Hungary, 2005, pp. 1--5



Interpolando, extrapolando y comparando las curvas de acumulaci\'on de especies basadas en su incidencia


Robert K Colwell, Xuan Mao Chang, Chang Jing

Sobre diversidad biol{\'o}gica: el significado de las diversidades alfa, beta y gamma, GORFI, 2005, pp. 73--84



Un nuevo m\'etodo estad\'\istico para la evaluaci\'on de la similitud en la composici\'on de especies con datos de incidencia y abundancia


Robert K Colwell, Anne Chao, Robin L Chazdon, Tsung-Jen Shen

Sobre diversidad biol{\'o}gica: el significado de las diversidades alfa, beta y gamma, GORFI, 2005, pp. 85--96



The Mid-Domain Effect: there’s a baby in the bathwater


Robert K Colwell, Carsten Rahbek, Nicholas J Gotelli

The American Naturalist, vol. 166, 2005, pp. E149--E154



Estimation of species richness: mixture models, the role of rare species, and inferential challenges


Chang Xuan Mao, Robert K Colwell

Ecology, vol. 86, Ecological Society of America, 2005, pp. 1143--1153



The influence of band sum area, domain extent, and range sizes on the latitudinal mid-domain effect


Tom S Romdal, Robert K Colwell, Carsten Rahbek

Ecology, vol. 86, Ecological Society of America, 2005, pp. 235--244



Estimating the species accumulation curve using mixtures


Chang Xuan Mao, Robert K Colwell, Jing Chang

Biometrics, vol. 61, Blackwell Publishing 350 Main Street, Malden, MA 02148, USA, and PO Box 1354~…, 2005, pp. 433--441



Arachnida


Jonathan A Coddington, G Giribet, MS Harvey, L Prendini, DE Walter

Assembling the tree of life, Oxford University Press, 2004



Interpolating, extrapolating, and comparing incidence-based species accumulation curves


Robert K Colwell, Chang Xuan Mao, Jing Chang

Ecology, vol. 85, Ecological Society of America, 2004, pp. 2717--2727



The mid-domain effect and species richness patterns: What have we learned so far?


Robert K Colwell, Carsten Rahbek, Nicholas J Gotelli

The American Naturalist, vol. 163, 2004, pp. E1--E23



The coincidence of rarity and richness and the potential signature of history in centres of endemism


Walter Jetz, Carsten Rahbek, Robert K Colwell

Ecology Letters, vol. 7, Blackwell Science Ltd Oxford, UK, 2004, pp. 1180--1191



Species coextinctions and the biodiversity crisis


Lian Pin Koh, Robert R Dunn, Navjot S Sodhi, Robert K Colwell, Heather C Proctor, Vincent S Smith

science, vol. 305, American Association for the Advancement of Science, 2004, pp. 1632--1634



Wings and stings


R. K. Colwell, J. T. Longino

Natural History, vol. 112, American Museum of Natural History, 2003, pp. 13--13



The ant fauna of a tropical rain forest: estimating species richness three different ways


John T Longino, Jonathan Coddington, Robert K Colwell

Ecology, vol. 83, Ecological Society of America, 2002, pp. 689--702



An evolutionary perspective on human interactions with biodiversity.


RK Colwell

AMERICAN JOURNAL OF HUMAN BIOLOGY, vol. 13, WILEY-LISS DIV JOHN WILEY \& SONS INC, 605 THIRD AVE, NEW YORK, NY 10158-0012 USA, 2001, pp. 116--117



Quantifying biodiversity: procedures and pitfalls in the measurement and comparison of species richness


Nicholas J Gotelli, Robert K Colwell

Ecology letters, vol. 4, Wiley Online Library, 2001, pp. 379--391



The mid-domain effect and the longitudinal dimension of continents-Reply


R Colwell, D Lees

Trends in Ecology \& Evolution, vol. 15, ELSEVIER SCIENCE LONDON 84 THEOBALDS RD, LONDON WC1X 8RR, ENGLAND, 2000, pp. 289--289



Rensch’s rule crosses the line: convergent allometry of sexual size dimorphism in hummingbirds and flower mites


Robert K Colwell

The American Naturalist, vol. 156, The University of Chicago Press, 2000, pp. 495--510



A barrier runs through it… or maybe just a river


Robert K Colwell

Proceedings of the National Academy of Sciences, vol. 97, National Academy of Sciences, 2000, pp. 13470--13472



Hummingbirds and flower mites: contrasts and conflicts between two nectarivores


Robert K Colwell

A ornitologia no Brasil: pesquisa atual e perspectivas. Anais do VII Congresso Brasileiro de Ornitologia, Rio de Janeiro, 2000, pp. 65--91



The mid-domain effect: geometric constraints on the geography of species richness


Robert K Colwell, David C Lees

Trends in ecology \& evolution, vol. 15, Elsevier Current Trends, 2000, pp. 70--76



Reply from RK Colwell and DC Lees


Robert Colwell, David Lees

Trends in Ecology \& Evolution, vol. 15, Elsevier, 2000, p. 289



Т he extent, distribution, and biology of invertebrate


IAN OLIVER, ANTHONY PIK, DAVID BRITTON, J MARK DANGERFIELD, ROBERT K COLWELL

BioScience, vol. 50, 2000, pp. 441--450



Virtual biodiversity assessment systems


Ian Oliver, Anthony Pik, David Britton, J Mark Dangerfield, Robert K Colwell, Andrew J Beattie

Bioscience, vol. 50, American Institute of Biological Sciences Circulation, AIBS, 1313 Dolley~…, 2000, pp. 441--450



Tropical tree richness and resource-based niches


Robin L Chazdon, Robert K Colwell, Julie S Denslow

Science, vol. 285, American Association for the Advancement of Science, 1999, pp. 1459--1459



The evolution of host-plant affiliation, body size and sexual dimorphism in hummingbird flower mites (Ascidae: Rhinoseius and Tropicoseius)


RK Colwell, P Naskrecki

In G. R. Needham, R. Mitchell, D. J. Horn, and W. C. Welbourn, editors. Acarology IX, Volume 2: Symposia (IXth International Congress of Acarology ), Ohio Biological Survey, Columbus, Ohio, 1999, pp. 79--86



Sexual sorting in hummingbird flower mites (Mesostigmata: Ascidae)


Robert K Colwell, Shadid Naeem

Annals of the Entomological Society of America, vol. 92, Oxford University Press Oxford, UK, 1999, pp. 952-959



Biodiversity assessment using structured inventory: capturing the ant fauna of a tropical rain forest


John T Longino, Robert K Colwell

Ecological applications, vol. 7, Ecological Society of America, 1997, pp. 1263--1277



The Flower Mites of Trinidad: The Genus Rhinoseius (Acari: Ascidae)


Barry M OConnor, Robert Knight Colwell, Shahid Naeem

Museum of Zoology, University of Michigan, 1997



Integration of species and ecosystem approaches to conservation


SL Tartowski, EB Allen, NE Barrett, AR Berkowitz, RK Colwell

S. T. A. Pickett, R. S. Ostfield, M. Shachak, and G. E. Likens, editors. Enhancing the ecological basis of conservation: Heterogeneity, ecoystem function, and biodiversity. Proceedings of the Sixth Cary Conference, Institute of Ecosystem Studies, 1997, pp. 187--198



Toward a resolution of conflicting paradigms


SL Tartowski, EB Allen, NE Barrett, AR Berkowitz, Robert K Colwell, PM Groffman, J Harte, Hugh Philip Possingham, CM Pringe, DL Strayer, others

Chapman \& Hall, 1997



Quantitative inventory of a lowland rainforest ant fauna. Inventario cuantitativo de la fauna de hormigas en un bosque lluvioso de las tierras bajas.


John T Longino, Robert K Colwell

Bulletin of the Ecological Society of America., vol. 77, 1996, p. 271



New genus and two new species of Melicharini from Venezuela (Acari: Mesostigmata: Ascidae)


Piotr Naskrecki, Robert K Colwell

Annals of the Entomological Society of America, vol. 88, Oxford University Press Oxford, UK, 1995, pp. 284--293



Excelsotarsonemus kaliszewskii, a new genus and new species from Costa Rica (Acari: Tarsonemidae)


Ronald Ochoa, Piotr Naskrecki, Robert K Colwell

International journal of acarology, vol. 21, Taylor \& Francis Group, 1995, pp. 67--74



Potential ecological and evolutionary problems of introducing transgenic crops into the environment


Robert K Colwell

biosafety for sustainable agriculture: Sharing biotechnology regulatory experiences of the western hemisphere. Eds. Krattiger, AF \& Rosemarin A. ISAAA, Ithaca \& SEI, Stockholm, 1994, pp. 33--46



Human aspects of biodiversity: An evolutionary perspective


Robert K Colwell

Biodiversity and Global Change, Cab International Wallingford, 1994, pp. 211--24



Estimating terrestrial biodiversity through extrapolation


Robert K Colwell, Jonathan A Coddington

Philosophical Transactions of the Royal Society of London. Series B: Biological Sciences, vol. 345, The Royal Society London, 1994, pp. 101--118



Nonbiological gradients in species richness and a spurious Rapoport effect


Robert K Colwell, George C Hurtt

The American Naturalist, vol. 144, University of Chicago Press, 1994, pp. 570--595



Life-history patterns of hummingbird flower mites in relation to host phenology and morphology


Robert K Colwell, Shahid Naeem

Mites, Springer, Boston, MA, 1994, pp. 23-44



Diversity of long-term flowering patterns


LE Newstrom, GW Frankie, HG Baker, RK Colwell

In L. A. McDade, K. S. Bawa, H. A. Hespenheide, and G. S. Hartshorn, editors. La Selva: Ecology and natural history of a Neotropical rain forest., University of Chicago Press, 1994, pp. 142--160



Terrestrial arthropod assemblages: their use in conservation planning


C Kremen, RK Colwell, TL Erwin, DD Murphy, RF Noss, MA and Sanjayan

Conservation biology, Blackwell Scientific Publications, 1993, pp. 796--808



Cellulose acetate electrophoretic techniques for the genetic analysis of individual ascid mites (Mesostigmata: Ascidae)


Kim M Christiansen, Robert K Colwell, Marek J Kaliszewski

International journal of acarology, vol. 18, Taylor \& Francis, 1992, pp. 97--105



Niche: a bifurcation in the conceptual lineage of the term


Robert K Colwell

Keywords in evolutionary biology, Harvard University press, MA, USA, 1992, pp. 241--248



Making Sense of Ecological Complexity: A Personal and Conceptual Restrospective


Robert K Colwell

Biotropica, Association for Tropical Biology, 1992, pp. 226--232



The polymerase chain reaction and sequencing of mite DNA


MJ Kaliszewski, J Tobolewski, S Seyoum, I Chojnacki, MM Kaliszewska, DJ Stanton, RK Colwell

International journal of acarology, vol. 18, Taylor \& Francis, 1992, pp. 231--239



Detection and identification of mammalian DNA from the gut of museum specimens of ticks


Jerzy Tobolewski, Marek J Kaliszewski, Robert K Colwell, James H Oliver Jr

Journal of medical entomology, vol. 29, Oxford University Press Oxford, UK, 1992, pp. 1049--1051



Conceptual framework and research issues for species diversity at the community level


R Barbault, RK Colwell, B Dias, DL Hawksworth, M Huston, P Laserre, D Stone, T Younes

From genes to ecosystems: a research agenda for biodiversity. IUBS, Cambridge, Mass, 1991, pp. 37--71



Host plant discrimination: Experiments with hummingbird flower mites


AJ Heyneman, Robert K Colwell, S Naeem, DS Dobkin, B Hallet

Plant-animal interactions: evolutionary ecology in tropical and temperate regions. Price, P. W. Lewinsohn, T. M. Fernandes, G. W. Benson, W. W., Eds., John Wiley and Sons, 1991, pp. 455--485



Ecological consequences of heterogeneity of consumable resources


Shahid Naeem, Robert K Colwell

Ecological heterogeneity, Springer, New York, NY, 1991, pp. 224--255



Flower mites of Trinidad II. The genus Proctolaelaps (Acari: Ascidae)


Barry M OConnor, Robert K Colwell, Shahid Naeem

The Great Basin Naturalist, Brigham Young University, 1991, pp. 348--376



Hummingbirds of the Juan Fern\'andez Islands: natural history, evolution and population status


RK Colwell

Ibis, vol. 131, Blackwell Publishing Ltd (NOTE: that the names of the birds in the Abstract~…, 1989, pp. 548--566



Hummingbirds of the Juan Fern\'andez Islands: natural history, evolution and population status


RK Colwell

Ibis, vol. 131, Blackwell Publishing Ltd (NOTE: that the names of the birds in the Abstract~…, 1989, pp. 548--566



Natural and unnatural history: Biological diversity and genetic engineering. Pages 1-40 in W. R. Shea and B. Sitter, editors. Scientists and their responsibility. Watson International Publishing, Canton, MA.


Robert K Colwell

W. R. Shea and B. Sitter, editors. Scientists and their responsibility., Watson International Publishing, Canton, MA., 1989, pp. 1--40



Natural and unnatural history: Biological diversity and genetic engineering. Pages 1-40 in W. R. Shea and B. Sitter, editors. Scientists and their responsibility. Watson International Publishing, Canton, MA.


Robert K Colwell

W. R. Shea and B. Sitter, editors. Scientists and their responsibility., Watson International Publishing, Canton, MA., 1989, pp. 1--40



Hutchinsonian ratios and statistical power


Jonathan B Losos, Shahid Naeem, Robert K Colwell

Evolution, vol. 43, Society for the Study of Evolution, Wiley, 1989, pp. 1820--1826



Another reading of NAS gene report


Robert K Colwell

BioScience;(United States), vol. 38, Univ. of Tennessee, Knoxville (USA), 1988



Ecology and biotechnology: expectations and outliers


Robert K Colwell

NATO ASI Series. Series G: Ecological Sciences, vol. 18, Plenum, 1988, pp. 163--180



Another reading of NAS gene report


Robert K Colwell

BioScience;(United States), vol. 38, Univ. of Tennessee, Knoxville (USA), 1988



Response to the Office of Science and Technology Policy Notice" Coordinated Framework for Regulation of Biotechnology"


Robert K Colwell, Lawrence W Barnthouse, Andrew Dobson, Frieda Taub, Richard Wetzler

Bulletin of the Ecological Society of America, vol. 68, Wiley, Ecological Society of America, 1987, pp. 16--23



Population structure and sexual selection for host fidelity in the speciation of hummingbird flower mites


RK Colwell

Evolutionary processes and theory. Karlin, S, and E. Nevo, Eds., Academic, New York, New York, USA, 1986, pp. 475--495



Community biology and sexual selection: Lessons from hummingbird flower mites


Robert K Colwell

Community Ecology. Diamond, J.; Case, TJ (eds.)., Harper and Row, N. Y., 1986, pp. 406-424



The evolution of ecology


Robert K Colwell

American Zoologist, vol. 25, Oxford University Press, 1985, pp. 771--777



Stowaways on the Hummingbird Express


Robert K Colwell

Natural History, vol. 94, 1985, pp. 56--63



Genetic engineering in agriculture


Robert K Colwell, Elliott A Norse, David Pimentel, Frances E Sharples, Daniel Simberloff

Science, vol. 229, American Association for the Advancement of Science, 1985, pp. 111--112



Report on workshop on possible ecological and evolutionary impacts of bioengineered organisms released into the environment


James H Brown, Robert K Colwell, Richard E Lenski, Bruce R Levin, Monte Lloyd, Philip J Regal, Daniel Simberloff

Bulletin of the Ecological Society of America, vol. 65, Wiley, Ecological Society of America, 1984, pp. 436--438



A null model for null models in biogeography


R. K. Colwell, D. W. Winkler

Jr. Strong D. R., D. Simberloff, L. G. Abele, A. B. Thistle, Ecological communities: Conceptual issues and the evidence, chapter 20, Princeton University Press, 1984, pp. 344-359



Comparing behavior: Studying man studying animals


RK Colwell, MC King, DW Rajecki

1983



Theoretical Ecology: Principles and Applications


Robert K Colwell

American Ornithological Society, 1983



CITATION CLASSIC-ON THE MEASUREMENT OF NICHE BREADTH AND OVERLAP


RK Colwell

CURRENT CONTENTS/AGRICULTURE BIOLOGY \& ENVIRONMENTAL SCIENCES, INST SCI INFORM INC 3501 MARKET ST, PHILADELPHIA, PA 19104, 1982, pp. 18--18



Female-biased sex ratios (reply)


Robert K Colwell

Nature, vol. 298, Nature Publishing Group, 1982, pp. 495--496



The geographical ecology of hummingbird flower mites in relation, to their host plants and carriers.


Robert K Colwell

Recent Advances in Acarology., vol. 2, 1979, pp. 461--468



Elevation and the morphology, flight energetics, and foraging ecology of tropical hummingbirds


Peter Feinsinger, Robert K Colwell, John Terborgh, Susan Budd Chaplin

The American Naturalist, vol. 113, University of Chicago Press, 1979, pp. 481--497



EXPERIMENTAL STUDIES OF THE NICHE


R. K. Colwell, E. Fuentes

1975



Predictability, constancy, and contingency of periodic phenomena


Robert K Colwell

Ecology, vol. 55, Ecological Society of America, 1974, pp. 1148--1153



Competition for the nectar of Centropogon valerii by the hummingbird Colibri thalassinus and the flower-piercer Diglossa plumbea, and its evolutionary implications


Robert K Colwell, Burr J Betts, Pille Bunnell, F Lynn Carpenter, Peter Feinsinger

The condor, vol. 76, American Ornithological Society, 1974, pp. 447--452



Competition and coexistence in a simple tropical community


Robert K Colwell

The American Naturalist, vol. 107, University of Chicago Press, 1973, pp. 737-760



Ecological Assessment


Matthew H Greenstone, Robert K Colwell, Frank A Pitelka

Science, vol. 177, American Association for the Advancement of Science, 1972, pp. 210--210



On the measurement of niche breadth and overlap


Robert K Colwell, Douglas J Futuyma

Ecology, vol. 52, Ecological Society of America, 1971, pp. 567--576



The relationship between species diversity and stability: an experimental approach with protozoa and bacteria


NG Hairston, JD Allan, RK Colwell, DJ Futuyma, J Howell, MD Lubin, J Mathias, JH Vandermeer

Ecology, vol. 49, Ecological Society of America, 1968, pp. 1091--1101


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